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I think it has to do with what happened in the Ashkenazi diaspora. All through the Middle Ages, the Jews were universally well educated and knew how to read and converse in Hebrew. In the Central and Eastern Europe, especially after the Exile from Spain, the fate of the Jews was not as simple. In the villages and shtetls, education had to yield to survival. In the cities, Jewish studies had to yield to European education: in order to survive, the Jews had to learn to communicate with the host nations. In order to advance up from the mere survival level, the Jews had to become the best in commerce, finance, medicine, sciences, music, literature, agriculture, etc. There are only 24 hours in a day, even for a Jew. Result - Jewish education started to fall behind. There stared to appear cases when a Cohen or a Levite could not read a text from Tanach. What to do? It had to be whoever was the most educated in the Kehillah. Typically the most educated were those whose parents had accumulated enough wealth that their children could afford going to yeshivoth - or those whose parents could not afford to educate their children in the non-Jewish studies and therefore were educated by the community.
So, there was a strange split occurring in the Jewish society right around the 16-19th century: a reduction in the proportion of the Leviim and Cohanim who were were educated in Jewish subjects, accompanied by an increase in the proportion of educated "common" Jews. So more an more common was a situation when the Cohain or Levite was called for aliyah, all he could read or even memorize was just the brochah.
Such man got very little respect from the congregation. Davening and reading Torah started to become a formality, like in the times of Isaiah. The only way possible to keep the congregation was to allow its most educated members to read Torah, even if they were not a Cohain or Levi. At the early onset of this trend, they would announce the name of the Cohen or Levite in whose stead they were reading. Later even that went away, and today in most shuls it is merely the most respected member of the congregation gets to perform the aliyah.
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